Asplenium septentrionale is a fern (rhizomatous) that is native to California, and also found elsewhere in North America and beyond.
California Rare Plant Rank: 2B.3 (rare, threatened, or endangered in CA; common elsewhere).
[Wikipedia] Hybridizations: Asplenium septentrionale can hybridize with maidenhair spleenwort (A. trichomanes) to form the hybrid A. × alternifolium, with A. foreziense to form A. × costei, with black spleenwort (A. adiantum-nigrum) to form A. × contrei, with A. obovatum subsp. lanceolatum to form A. × souchei, and with wall-rue (A. ruta-muraria) to form A. × murbeckii and A. × tavelio. A. × murbeckii can back-cross with A. septentrionale to form A. × hungaricum. (link added by Mary Ann Machi)
Calflora:
Information on California plants for education, research and conservation,
with data contributed by
public and private institutions and individuals.
[web application]. 2025.The Calflora Database
[a non-profit organization].Available:
https://www.calflora.org/(Accessed: 03/28/2025).